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  • Blue American bullfrog, Rana catesbeiana, a rare mutation of a normally green frog caused by a lack of yellow pigment in the frog's skin. Mendocino County, California.
    13090017.jpg
  • Students Zito Afranio (left), Marianna Tucci, Benny Carvalho, and Scott Heacox (back to camera) search for lizards in the Liquica district of Timor-Leste (East Timor). They are participating in an ongoing survey of Timorese reptiles and amphibians.
    10041716-east-timor.jpg
  • Herpetologist Mark O'Shea and students Laca Ribeiro, Zito Afranio, Benny Carvalho, and Luis Lemos search for lizards in the Liquica district of Timor-Leste (East Timor). They are participating in an ongoing survey of Timorese reptiles and amphibians.
    10041715-east-timor.jpg
  • American students Caitlin Sanchez, Jester Ceballos, Marianna Tucci, and Eric Leatham are joined by Timorese Benny Carvalho in a search for lizards in a riverbed in the Liquica district of Timor-Leste (East Timor). They are participating in an ongoing survey of Timorese reptiles and amphibians.
    10041705-east-timor.jpg
  • Timorese student Laca Ribeiro captures a tokay gecko, Gekko gecko, in a tree in the Liquica district of Timor-Leste (East Timor). He is participating in an ongoing survey of Timorese reptiles and amphibians.
    10041686-east-timor.jpg
  • Timorese student Laca Ribeiro captures a tokay gecko, Gekko gecko, in a tree in the Liquica district of Timor-Leste (East Timor). He is participating in an ongoing survey of Timorese reptiles and amphibians.
    10041684-east-timor.jpg
  • Using his sock as a glove, Timorese student Laca Ribeiro captures a tokay gecko, Gekko gecko, in a tree in the Liquica district of Timor-Leste (East Timor), while Luis Lemos watches. They are participating in an ongoing survey of Timorese reptiles and amphibians.
    10041681-east-timor.jpg
  • Timorese student Laca Ribeiro searches for lizards in a tree in the Liquica district of Timor-Leste (East Timor). He is participating in an ongoing survey of Timorese reptiles and amphibians.
    10041663-east-timor.jpg
  • Timorese student Laca Ribeiro captures a Timorese flying dragon, Draco timoriensis, in a tree in the Liquica district of Timor-Leste (East Timor). He is participating in an ongoing survey of Timorese reptiles and amphibians.
    10041659-east-timor.jpg
  • Northern Pacific rattlesnake, Crotalus viridis oreganus, in a resting coil. Mount Diablo State Park, California
    07220074.jpg
  • Northern Pacific rattlesnake, Crotalus viridis oreganus, in a defensive posture, flicks out its tongue to "taste" the air with its Jacobson's organ. The snake's heat-sensing pit is visible below and in front of the eye. Mount Diablo State Park, California
    07220057.jpg
  • Red-eyed treefrog, Agalychnis callidryas. Villa Lapas, near Herradura, Costa Rica
    09062339.jpg
  • Red-eyed treefrog, Agalychnis callidryas. Villa Lapas, near Herradura, Costa Rica
    09062305.jpg
  • Red-eyed treefrog, Agalychnis callidryas. Villa Lapas, near Herradura, Costa Rica
    09062287.jpg
  • Red-eyed treefrog, Agalychnis callidryas. Villa Lapas, near Herradura, Costa Rica
    09062283.jpg
  • Cane toad, Bufo marinus. Captured in a fountain in front of Punta Leona hotel and resort, near Jaco, Costa Rica
    09062186-cane-toad.jpg
  • Cane toad, Bufo marinus. El General Valley, Costa Rica
    09063138-cane-toad.jpg
  • Black spiny-tailed iguana, Ctenosaura similis. Manuel Antonio National Park, Costa Rica
    09062378.jpg
  • Black spiny-tailed iguana, Ctenosaura similis, on the grounds of Punta Leona Hotel and Resort, Costa Rica
    09062139.jpg
  • Black spiny-tailed iguana, Ctenosaura similis, on the grounds of Punta Leona Hotel and Resort, Costa Rica
    09062136-black-iguana.jpg
  • Black spiny-tailed iguana, Ctenosaura similis, on the grounds of Punta Leona Hotel and Resort, Costa Rica
    09062132-black-iguana.jpg
  • Black spiny-tailed iguana, Ctenosaura similis, on the grounds of Punta Leona Hotel and Resort, Costa Rica
    09062124-black-iguana.jpg
  • Black spiny-tailed iguana, Ctenosaura similis, in a tree on the shore of the Tarcoles River, Costa Rica
    09061905-black-iguana.jpg
  • Black spiny-tailed iguana, Ctenosaura similis, in a tree on the shore of the Tarcoles River, Costa Rica
    09061901-black-iguana.jpg
  • Red-eared slider, Trachemys scripta elegans. Captive at Zoo Ave, a zoo near San Jose, Costa Rica
    09060489-red-eared-slider.jpg
  • British herpetologist Mark O'Shea photographs a dog-faced water snake, Cerberus rynchops, captured in a rice paddy in the Baucau District of Timor-Leste (East Timor).
    10040947.jpg
  • British herpetologist Mark O'Shea photographs a dog-faced water snake, Cerberus rynchops, captured in a rice paddy in the Baucau District of Timor-Leste (East Timor).
    10040944.jpg
  • American herpetologist Hinrich Kaiser and student Caitlin Sanchez prepare specimens at their makeshift research station at Bakhita Mission, Near Eraulo, Ermera District, Timor-Leste (East Timor)
    10040935.jpg
  • British herpetologist Mark O'Shea, holding a dog-faced water snake, Cerberus rynchops, speaks to American and Timorese students at their makeshift research station at Bakhita Mission, Near Eraulo, Ermera District, Timor-Leste (East Timor). Left to right: Caitlin Sanchez, Scott Heacox, Professor Hinrich Kaiser, Luis Lemos, Mark O'Shea, Benny Carvalho, Zito Afranio, Marianna Tucci.
    10040914.jpg
  • American herpetologist Hinrich Kaiser prepares specimens at his makeshift research station at Tua Koin Resort, Atauro Island, Timor-Leste (East Timor)
    10040903.jpg
  • Metamorphosing black-spined toad, Duttaphrynus melanostictus, on the photographer's index finger.  Toadlet has four legs and is able to breathe air and walk on land, but has not fully resorbed its tail.  Dili District, Timor-Leste (East Timor).
    10040660-toad.jpg
  • Metamorphosing black-spined toad, Duttaphrynus melanostictus. Toadlet has four legs and is able to breathe air and walk on land, but has not fully resorbed its tail. Photographed on a US dime to show its size. Dili District, Timor-Leste (East Timor).
    10040658-toad.jpg
  • Metamorphosing black-spined toad, Duttaphrynus melanostictus. Toadlet has four legs and is able to breathe air and walk on land, but has not fully resorbed its tail. Photographed on a US dime to show its size. Dili District, Timor-Leste (East Timor).
    10040656-toad.jpg
  • Metamorphosing black-spined toad, Duttaphrynus melanostictus. Toadlet has four legs and is able to breathe air and walk on land, but has not fully resorbed its tail. Photographed on a US dime to show its size. Dili District, Timor-Leste (East Timor).
    10040654-toad.jpg
  • Herpetologist Mark O'Shea photographs a Sunda Island pitviper, Cryptelytrops insularis. Baucau District of Timor-Leste (East Timor).
    10040592-pitviper.jpg
  • Timor flying dragon, Draco timoriensis.  A herpetologist is holding the lizard by its tail to show its "wings" -- flaps of skin stretched over modified ribs, which it uses to glide from high branches.  Liquica District, Timor-Leste (East Timor)
    10040590-flying-dragon.jpg
  • Dog-faced water snake or bockadam, Cerberus rynchops. An aquatic species, it spends most of its time partially or completely submerged in flooded rice paddies; this one was moved onto the grass to be photographed. Baucau district, Timor-Leste (East Timor).
    10040555-water-snake.jpg
  • A previously undescribed species of night skink, tentatively called Eremiascincus sp. A, from Ermera Province, Timor-Leste (East Timor).
    10040398-skink.jpg
  • Common sun skink, Eutropis cf. multifasciata, near Eraulo, Ermera District, Timor-Leste (East Timor)
    10040370-skink.jpg
  • Tokay gecko, Gekko gecko, on Atauro Island, Timor-Leste (East Timor)
    10040347-gecko.jpg
  • Tokay gecko, Gekko gecko, on Atauro Island, Timor-Leste (East Timor)
    10040343-gecko.jpg
  • Tokay gecko, Gekko gecko, on Atauro Island, Timor-Leste (East Timor)
    10040339-gecko.jpg
  • Four-clawed gecko, Gehyra mutilata, on Atauro Island, Timor-Leste (East Timor).
    10040323-gecko.jpg
  • Four-clawed gecko, Gehyra mutilata, on Atauro Island, Timor-Leste (East Timor).
    10040305-gecko.jpg
  • Timor bent-toed gecko, a previously undescribed species of the genus Cyrtodactylus, from Atauro Island, Timor-Leste (East Timor).
    10040284-gecko.jpg
  • Timor bent-toed gecko, a previously undescribed species of the genus Cyrtodactylus, using its tongue to clean its eye.  Atauro Island, Timor-Leste (East Timor).
    10040270-gecko.jpg
  • Emerald tree skink, Lamprolepis cf smaragdina, from the Baucau district of Timor-Leste (East Timor).  While similar to L. smaragdina on other islands, individuals from East Timor are different in significant ways and will probably be assigned to a new species after further study.
    10040253-skink.jpg
  • Striped treefrog, Polypedates leucomystax, from the Baucau District of Timor-Leste (East Timor). This frog has only recently been studied in Timor-Leste, and will probably be assigned to another species.
    10040165-treefrog.jpg
  • Female rice paddy frog, Fejervarya sp., from the Baucau district of Timor-Leste (East Timor)
    10040089-rice-paddy-frog.jpg
  • Green frog, Rana clamitans melanota.  Native to eastern United States.
    05150245-green-frog.jpg
  • Green frog, Rana clamitans melanota.  Native to eastern United States.
    05150241-green-frog.jpg
  • Tadpole of American bullfrog, Rana catesbeiana.  Ten days after hatching, the tadpole is about 8mm long. The skin is semi-transparent.
    05150200-bullfrog-tadpole.jpg
  • Tadpole of American bullfrog, Rana catesbeiana.  A week after hatching, the tadpole is about 8mm long and has developed the familiar head-and-tail shape. The skin is semi-transparent, and internal organs are visible.
    05150193-bullfrog-tadpole.jpg
  • Tadpole of American bullfrog, Rana catesbeiana.  Immediately after hatching, the tadpole is about 5mm long and still has the appearance of an embryo.
    05150098-bullfrog-tadpole.jpg
  • Pacific treefrog (Pacific chorus frog), Hyla regilla (Pseudacris regilla). Mount Diablo State Park, California
    05020265-pacific-treefrog.jpg
  • Pacific treefrog (Pacific chorus frog), Hyla regilla (Pseudacris regilla). Mount Diablo State Park, California
    05020263-pacific-treefrog.jpg
  • Pacific treefrog (Pacific chorus frog), Hyla regilla (Pseudacris regilla). Mount Diablo State Park, California
    05020153-pacific-treefrog.jpg
  • Pacific treefrog (Pacific chorus frog), Hyla regilla (Pseudacris regilla). Mount Diablo State Park, California
    05020149-pacific-treefrog.jpg
  • Pacific treefrog (Pacific chorus frog), Hyla regilla (Pseudacris regilla). Mount Diablo State Park, California
    05020142-pacific-treefrog.jpg
  • Female Yosemite toad, Bufo canorus, swimming in shallow water in a high elevation meadow in California's Sierra Nevada mountains. Also in the water are tadpoles of the same species. Once widespread in the high Sierra, since the 1980s Yosemite toads have disappeared or severely declined in over 50% of their former range, and now survive only in a handful of wet meadows. The toad is listed as a Species of Special Concern under the Federal Endangered Species Act as well as by the State of California, and is listed as Endangered by the IUCN.
    08150286.jpg
  • Blue American bullfrog, Rana catesbeiana, a rare mutation of a normally green frog caused by a lack of yellow pigment in the frog's skin. Mendocino County, California.
    13090009.jpg
  • Timorese student Laca Ribeiro searches for lizards in a tree in the Liquica district of Timor-Leste (East Timor). He is participating in an ongoing survey of Timorese reptiles and amphibians.
    10041667-east-timor.jpg
  • Timorese student Laca Ribeiro searches for lizards in a tree in the Liquica district of Timor-Leste (East Timor). He is participating in an ongoing survey of Timorese reptiles and amphibians.
    10041666-east-timor.jpg
  • British herpetologist Mark O'Shea and Timorese student Zito Afranio  search for lizards in a tree in the Liquica district of Timor-Leste (East Timor). They are participating in an ongoing survey of Timorese reptiles and amphibians.
    10041653-east-timor.jpg
  • Red-eyed treefrog, Agalychnis callidryas. Villa Lapas, near Herradura, Costa Rica
    09062349.jpg
  • Red-eyed treefrog, Agalychnis callidryas. Villa Lapas, near Herradura, Costa Rica
    09062327.jpg
  • Cane toad, Bufo marinus. Captured in a fountain in front of Punta Leona hotel and resort, near Jaco, Costa Rica
    09062191-cane-toad.jpg
  • Cane toad, Bufo marinus. Captured in a fountain in front of Punta Leona hotel and resort, near Jaco, Costa Rica
    09062190-cane-toad.jpg
  • Desert tortoise, Gopherus agassizi (Xerobates agassizi), surrounded by goldfields, Lasthenia chrysostoma. State- and Federally-listed Threatened Species. Desert Tortoise Natural Area, Mojave Desert, California.
    r296-desert-tortoise.jpg
  • Western fence lizard, Sceloporus occidentalis, Mount Diablo State Park, California
    09100027-fence-lizard.jpg
  • Western fence lizard, Sceloporus occidentalis, Mount Diablo State Park, California
    09090013-fence-lizard.jpg
  • Photographers Patricia Prettie (left) and Yamil Saenz approach a black spiny-tailed iguana, Ctenosaura similis, on the grounds of Punta Leona Hotel and Resort, Costa Rica
    09062116-black-iguana.jpg
  • British herpetologist Mark O'Shea milks the venom of a Sunda Island pitviper, Cryptelytrops insularis, at Bakhita Mission, near Eraulo, Ermera District, Timor-Leste (East Timor)
    10040989.jpg
  • British herpetologist Mark O'Shea prepares to milk the venom of a Sunda Island pitviper, Cryptelytrops insularis, at Bakhita Mission, near Eraulo, Ermera District, Timor-Leste (East Timor)
    10040987.jpg
  • British herpetologist Mark O'Shea photographs a dog-faced water snake, Cerberus rynchops, in a meadow near Eraulo, in the Ermera district of Timor-Leste (East Timor). The snake was originally captured in a rice paddy in the Baucau District.
    10040962.jpg
  • British herpetologist Mark O'Shea photographs a dog-faced water snake, Cerberus rynchops, in a meadow near Eraulo, in the Ermera district of Timor-Leste (East Timor). The snake was originally captured in a rice paddy in the Baucau District.
    10040950.jpg
  • Metamorphosing black-spined toad, Duttaphrynus melanostictus, on the photographer's index finger.  Toadlet has four legs and is able to breathe air and walk on land, but has not fully resorbed its tail.  Dili District, Timor-Leste (East Timor).
    10040659-toad.jpg
  • Tadpole of black-spined toad, Duttaphrynus melanostictus, with hind legs.  Dili District, Timor-Leste (East Timor).
    10040641-tadpole.jpg
  • Sunda Island pitviper, Cryptelytrops insularis, in defensive posture. The snake's heat-sensing pits are visible between the eye and the nostril. Cosidered an arboreal species throughout most of its range, but in Timor-Leste it is most often found on the ground. Baucau District of Timor-Leste (East Timor).
    10040616-pitviper.jpg
  • Herpetologist Mark O'Shea shows the fangs of a Sunda Island pitviper, Cryptelytrops insularis. Baucau District of Timor-Leste (East Timor).
    10040593-pitviper.jpg
  • Timor flying dragon, Draco timoriensis, a gliding lizard with "wings" -- flaps of skin stretched over modified ribs.  Liquica District, Timor-Leste (East Timor)
    10040572-flying-dragon.jpg
  • Dog-faced water snake or bockadam, Cerberus rynchops. An aquatic species, it spends most of its time partially or completely submerged in flooded rice paddies; this one was moved onto the grass to be photographed. Baucau district, Timor-Leste (East Timor).
    10040552-water-snake.jpg
  • Common sun skink, Eutropis cf. multifasciata, near Eraulo, Ermera District, Timor-Leste (East Timor)
    10040379-skink.jpg
  • Common sun skink, Eutropis cf. multifasciata, near Eraulo, Ermera District, Timor-Leste (East Timor)
    10040373-skink.jpg
  • Tokay gecko, Gekko gecko, on Atauro Island, Timor-Leste (East Timor)
    10040345-gecko.jpg
  • Emerald tree skink, Lamprolepis cf smaragdina, from the Baucau district of Timor-Leste (East Timor).  While similar to L. smaragdina on other islands, individuals from East Timor are different in significant ways and will probably be assigned to a new species after further study.
    10040259-skink.jpg
  • Four-fingered skink, Carlia sp., from the Baucau district of Timor-Leste (East Timor). A previously undescribed species, tentatively called Carlia species 3.  (Specimen HK548)
    10040208-skink.jpg
  • Striped treefrog, Polypedates leucomystax, from the Baucau District of Timor-Leste (East Timor). This frog has only recently been studied in Timor-Leste, and will probably be assigned to another species.
    10040171-treefrog.jpg
  • Striped treefrog, Polypedates leucomystax, from the Baucau District of Timor-Leste (East Timor). This frog has only recently been studied in Timor-Leste, and will probably be assigned to another species.
    10040163-treefrog.jpg
  • Hybrid African clawed frog, Silurana sp.  Native to sub-Saharan Africa, clawed frogs were exported worldwide in the 1940s and 1950s for use in human pregnancy tests, and were later sold widely as pets.  Feral populations are now established in many countries, including the United States, where they are a threat to native amphibians and fish.  Especially large populations exist in California, preying on endangered California red-legged frogs, Rana aurora draytonii, and endangered unarmored threespine sticklebacks, Gasterosteus aculeatus williamson.  Recent evidence suggests the chytrid fungus that is currently decimating frog populations around the world originated in Africa and was spread by these frogs.
    06050252-african-clawed-frog.jpg
  • Green frog, Rana clamitans melanota.  Native to eastern United States.
    05150247-green-frog.jpg
  • Female Yosemite toad, Bufo canorus, in a high elevation meadow in California's Sierra Nevada mountains. Once widespread in the high Sierra, since the 1980s Yosemite toads have disappeared or severely declined in over 50% of their former range, and now survive only in a handful of wet meadows. The toad is listed as a Species of Special Concern under the Federal Endangered Species Act as well as by the State of California, and is listed as Endangered by the IUCN.
    08150195.jpg
  • Female Yosemite toad, Bufo canorus, in a high elevation meadow in California's Sierra Nevada mountains. Once widespread in the high Sierra, since the 1980s Yosemite toads have disappeared or severely declined in over 50% of their former range, and now survive only in a handful of wet meadows. The toad is listed as a Species of Special Concern under the Federal Endangered Species Act as well as by the State of California, and is listed as Endangered by the IUCN.
    08150243.jpg
  • Female Yosemite toad, Bufo canorus, in a high elevation meadow in California's Sierra Nevada mountains. Once widespread in the high Sierra, since the 1980s Yosemite toads have disappeared or severely declined in over 50% of their former range, and now survive only in a handful of wet meadows. The toad is listed as a Species of Special Concern under the Federal Endangered Species Act as well as by the State of California, and is listed as Endangered by the IUCN.
    08150256.jpg
  • Female Yosemite toad, Bufo canorus, in a high elevation meadow in California's Sierra Nevada mountains. Once widespread in the high Sierra, since the 1980s Yosemite toads have disappeared or severely declined in over 50% of their former range, and now survive only in a handful of wet meadows. The toad is listed as a Species of Special Concern under the Federal Endangered Species Act as well as by the State of California, and is listed as Endangered by the IUCN.
    08150264.jpg
  • Herpetologist Mark O'Shea and students Laca Ribeiro, Marianna Tucci, Zito Afranio, Benny Carvalho, and Luis Lemos search for lizards in the Liquica district of Timor-Leste (East Timor). They are participating in an ongoing survey of Timorese reptiles and amphibians.
    10041714-east-timor.jpg
  • Timorese student Laca Ribeiro captures a tokay gecko, Gekko gecko, in a tree in the Liquica district of Timor-Leste (East Timor). He is participating in an ongoing survey of Timorese reptiles and amphibians.
    10041677-east-timor.jpg
  • Northern Pacific rattlesnake, Crotalus viridis oreganus, in a resting coil. The snake's heat-sensing pit is visible below and in front of the eye. Mount Diablo State Park, California
    07220070.jpg
  • Northern Pacific rattlesnake, Crotalus viridis oreganus, in a defensive posture, flicks out its tongue to "taste" the air with its Jacobson's organ. The snake's heat-sensing pit is visible below and in front of the eye. Mount Diablo State Park, California
    07220054.jpg
  • Red-eyed treefrog, Agalychnis callidryas. Villa Lapas, near Herradura, Costa Rica
    09062337.jpg
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